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Troublemaker – Stefano Unterthiner, Italy. Winner, animal portraits This young adult Sulawesi black-crested macaque, nicknamed Troublemaker, was more interested in the photographer than foraging for food, so getting a close-up wasn’t difficult. Troublemaker’s expression captures, Stefano says, "the spirit of these wonderful monkeys", and the setting makes it an unfor..

15–17 Years category winner: Yellowhammers fighting in winter by Fergus Gill (United Kingdom) Fergus lured the yellowhammers to his garden with sheaves of oats during the heavy snows last February.Photograph: Fergus Gill/Veolia Environnement Wildlife Photographer of the Year 2009

Animal Behaviour; All Other Animals category winner: Raindrop refresher by András Mészáros (Hungary)This is a category for photographs of animals that are not mammals or birds; in other words, the majority of animals on Earth. Most of them behave in ways that are seldom witnessed and little known or understood. After a summer shower, Andras headed off to his loc..

Underwater World – Beavering by Louis-Marie Préau (France)"I have never forgotten," says Préau, "the first time I saw a European beaver under water delivering a branch to its family." He is passionate about beavers and has returned each season for the past four years to the same spot on the Loire river in France to try to photograph what he saw that first time...

Behaviour: Mammals – Balancing Act by Joel Sartore (USA)In a death-defying manoeuvre, a mountain goat stretches to reach a mineral lick. Joel knew it was a favourite lick and had stationed himself at the other side of the gorge in Glacier National Park, Montana. When this female arrived, he watched as, slowly and methodically, she first balanced on all four feet on a sin..

Behaviour: Mammals – The Charge by Eric Pierre (France)Eric had been tracking Arctic wolves on Victoria Island, Canada, when his guide spotted a herd of muskoxen 3 miles (5km) away. Approaching, Eric could see that the herd was nervous, probably because wolves were also on its trail. He made a detour and stopped about half a mile away upwind. Suddenly, he realised that t..

Behaviour: Birds – Taking Flight, Paul GoldsteinPaul arrived very early on the shores of Lake Nakuru, Kenya, before the rising sun had burned off the mist. He had returned to photograph the greater and lesser flamingos and used shade, shadow and silhouette to create drama, rather than emphasising their vivid colours with sunlight. He was helped by a combination of circum..